Copyright notice: PDFs available for personal and educational use only; any further distribution, publication or commercial usage requires additional permission. Copyright and all rights retained by the authors and journals.

Book Chapters

Mills, S.C., Mokkonen, M., Koskela, E. & Mappes T. 2014 GEIs and reliable signaling of male quality in bank voles. In: Genotype-by-Environment Interactions and Sexual Selection Eds.: Hunt, J & Hosken, D. Wiley Blackwell, Oxford pdf

Lecchini, D., Mills, S.C. and R. Beldade 2015. Reproduction, Dispersal and Larval recruitment. In: Damselfishes Ed.: Parmentier, E. Taylor and Francis, New York.



55. Beldade, R., Blandin, A., O’Donnell, R. & Mills, S.C. 2017 It takes three to tango: Cascading fitness effects of anemone bleaching on associated anemonefish hormones and reproduction. Nature Communications in press

54. Nedelec, S.L., Mills, S.C., Radford, A.N., Beldade, R., Simpson, S.D., Nedelec, B. & Côté, I. 2017 Motorboat noise disrupts co-operative interspecific interactions. Scientific Reports in press

53. Nanninga, G.B., Côté, I.M., Beldade, R. & Mills, S.C. 2017. Behavioural acclimation to cameras and observers in coral reef fishes. Ethology in press

52. Armstrong, E.J, Allen, T., Beltrand, M., Dubousquet, V., Stillman, J.H. & Mills, S.C. 2017 Interactive effects of ocean acidification and warming on the development and physiology of a mollusc, the blue-ring sea hare, Stylocheilus striatus. Marine Biology 164: 107 pdf.

51. Bornancin, L., Bonnard, I., Mills, S.C. & Banaigs, B. 2017 Chemical mediation as a structuring element in marine gastropod predator-prey interactions. Natural Products Report pdf.

50. Horwitz, R., Jackson, M. & Mills, S.C. 2017. The embryonic life-history of the tropical sea hare Stylocheilus striatus (Gastropoda: Opisthobranchia) under ambient and elevated ocean temperatures. PeerJ 5:e2956; DOI 10.7717/peerj.2956 pdf

49. O’Donnell, J., Beldade, R., Mills, S.C., Williams, H. & Bernardi, G. 2017. Life history, larval dispersal, and connectivity in coral reef fish among the Scattered Islands of the Mozambique Channel. Coral Reefs 36: 223-232 pdf


48. Gil, M.A., Goldenberg, S., Thai Bach, A.L., Mills, S.C. & Claudet, J. 2016 Interactive effects of three pervasive marine stressors in a post-disturbance coral reef Coral Reefs 35: 1281-1293 pdf

47. Nedelec, S.L., Mills, S.C., Nedelec, B., Lecchini, D., Simpson, S.D. & Radford, A.N. 2016 Repeated exposure to noise increases tolerance in a coral reef fish. Environmental Pollution 216:428-436 pdf

46. Mokkonen, M., Koskela, E., Mappes, T. & Mills, S.C. 2016 Evolutionary conflict between maternal and paternal interests: Integration with evolutionary endocrinology. Integrative and Comparative Biology 56(2): 146-158 pdf

45. Bornancin, L., Boyaud, F., Mahiout, Z., Bonnard, I., Mills, S.C., Banaigs, B. & Inguimbert, N. 2016 Isolation and synthesis of laxaphycin B-type peptides: a case study and clues on their biosynthesis. Marine Drugs 13: 7285-7300 pdf


44. Moutardier G, Gereva S, Mills SC, Adjeroud M, Beldade R, Ham J, Kaku R & Dumas P. 2015 Lime juice and vinegar injections as a cheap and natural alternative to control COTS outbreaks. PLoS One 10(9): e0137605. doi:10.1371/journal.pone.0137605 pdf

43. Mills, S.C., Beldade., R Chabanet, P., Bigot, L., O'Donnell, J. & Bernardi, G. 2015. Ghosts of thermal past: reef fish exposed to historic high temperatures have heightened stress response to further stressors Coral Reefs 34: 1255-1260 pdf

42. Leray M, Meyer CP, Mills S.C. 2015. Metabarcoding dietary analysis of coral dwelling predatory fish demonstrates the minor contribution of coral mutualists to their highly partitioned, generalist diet. PeerJ 3:e1047 pdf

41. Beldade, R., Mills, S.C., Claudet, J. & I. Côté. 2015. More coral, more fish? Contrasting snapshots from a remote Pacific atoll. PeerJ 3:e745 pdf


40. Rouzé, H., Lecellier, G., Mills, S.C., Planes, S., Berteaux-Lecellier, V. & Stewart, H. 2014 Juvenile Trapezia spp. crabs can increase juvenile host coral survival by protection from predation Marine Ecology Progress Series 515: 151-159 pdf

39. Nedelec, S.L., Radford, A.N., Simpson, S.D., Nedelec, B., Lecchini. D, & Mills S.C. 2014 Anthropogenic noise playback impairs embryonic development and increases mortality in a marine invertebrate. Scientific Reports 4, 5891 pdf


38. Leray M., Yang J.Y., Meyer C.P., Mills S.C., Agudelo N.C., Ranwez V., Boehm J.T. & Machida R.J. 2013 A new versatile primer set targeting a short fragment of the mitochondrial COI region for metabarcoding metazoan diversity: application for characterizing coral reef fish gut contents Frontiers in Zoology 10: 34 pdf

37. Leray, M., Agudelo, N.C., Mills S.C., & Meyer, C.P. 2013 Effectiveness of annealing blocking primers versus restriction enzymes for characterization of generalist diets: unexpected prey revealed in the gut contents of two coral reef fish species. PLoS One 8(4): e58076 pdf

36. Mourier, J., Mills, S.C., Planes, S. 2013 Population structure, spatial distribution and life-history traits of blacktip reef sharks Carcharhinus melanopterus. Journal of Fish Biology 82: 979-993 pdf


35. Leray M., Beraud M., Anker A., Chancerelle Y. & Mills S.C. 2012 Acanthaster planci outbreak: Decline in coral health, coral size structure modification and consequences for obligate decapod assemblages. PLoS One 7(4): e35456 pdf

34. Stier, A.C., Gil, M.A., Seabird McKeon, C., Lemer, S., Leray, M., Mills S.C., & Osenberg, C.W. 2012 Housekeeping mutualisms: do more symbionts facilitate host performance? PLoS One 7(4): e32079 pdf

33. Mills S.C. 2012 Density-dependent prophylaxis in the coral-eating crown-of-thorns sea star, Acanthaster planci. Coral Reefs 31: 603-612 pdf

32. Leray, M., Boehm, J.T., Mills S.C., & Meyer, C.P. 2012 Moorea BIOCODE barcode library as a tool for understanding predator-prey interactions: insights into the diet of common predatory coral reef fishes. Coral Reefs 31(2): 383-388 pdf

31. Lecchini D., Brié C., Frédérich, B., Galzin, R., Nakamura, Y., Carassou, L. & Mills S.C. 2012 Effects of alternate reef states on coral reef fish habitat associations Environmental Biology of Fishes 94(2): 421-429 pdf

30. Mills S.C., Koskela, E. & Mappes, T. 2012 Intralocus sexual conflict for fitness: sexually antagonistic alleles for testosterone Proceedings of the Royal Society of Lond B 279: 1889-1895 pdf

29. Mokkonen, M., Koskela, E., Mappes, T. & Mills, S.C. 2012 Sexual antagonism for testosterone maintains multiple mating behaviour. Journal of Animal Ecology 81(1): 277-283 pdf

28. Mappes, T., Aspi, J., Koskela, E., Mills S.C., Poikonen, T. & Tuomi, J. 2012 Advantage of rare infanticide strategies in an invasion experiment of behavioural polymorphism. Nature Communications 3: 611 pdf

27. Santos R., Radford, C.A., Poupin, J., Brié C., Mills, S.C., Galzin, R. & Lecchini D. 2012 Temporal patterns in the post-larval supply of two crustacean taxa in Rangiroa Atoll, French Polynesia. Fisheries Science 78(1): 75-80 pdf


26. Mokkonen, M., Kokko, H., Koskela, E., Lehtonen, J., Mappes, T., Martiskainen, H. & Mills, S.C. 2011 Negative frequency-dependent selection of sexually antagonistic alleles in Myodes glareolus Science 334(6058): 972-974 pdf SupplMat

25. Lecchini D., Mills S.C., Brié C., Lo C. M., & B. Banaigs. 2011 Chemical stimuli in coral reefs: how butterflyfishes find their food Environmental Biology of Fishes 91(3): 303-309 pdf


24. Mills, S.C. & Côté, I. 2010 Crime and punishment in a roaming cleaner fish Proceedings of the Royal Society of London B 277(1700): 3617-3622 pdf

23. Frédérich, B., Mills, S.C., Denoël, M., Parmentier, E., Brié, C., Santos, R., Waqalevu, V.P. & Lecchini, D. 2010 Colour differentiation in a coral reef fish throughout ontogeny: habitat backgound and flexibility Aquatic Biology 9(3): 271-277

22. Schroderus, E., Jokinen, I., Koivula, M., Koskela, E., Mappes T., Mills S.C., Oskanen, T.A. & Poikonen, T. 2010 Intra- and intersexual trade-offs between testosterone and immune system: implications for sexual and sexually antagonistic selection American Naturalist 176(4): E90-E97 pdf

21. Mills, S.C., Mourier, J. & Galzin R. 2010 Plasma cortisol and 11-ketotestosterone enzyme immunoassay (EIA) kit validation for three fish species: the orange clownfish Amphiprion percula, the orangefin anemonefish Amphiprion chrysopterus and the blacktip reef shark Carcharhinus melanopterus Journal of Fish Biology 77: 769-777 pdf

20. Lecchini, D., Mills, S.C., Brié, C., Maurin, R., & Banaigs, B. 2010 Ecological determinants and sensory mechanisms in habitat selection of crustacean post-larvae Behavioral Ecology 21: 599-607 pdf

19. Mills S.C., Grapputo, A., Jokinen, I., Koskela, E., Mappes T. & Poikonen, T. 2010 Fitness trade-offs mediated by immunosuppression costs in a small mammal Evolution 64(1): 166-179 pdf


18. Planes, S., Lecchini, D., Romans, P., & Mills, S.C. 2009 Effects of post-settlement mortality on life history traits (size and parasite load) of Mediterranean juvenile fish (Diplodus vulgaris and Diplodus sargus) Aquatic Biology 6(1-3): 153-158 pdf

17. Mills S.C., Grapputo, A., I. Jokinen, Koskela, E., Mappes T., Oskanen, T.A. & Poikonen, T. 2009 Testosterone-mediated effects on fitness-related phenotypic traits and fitness American Naturalist 173(4): 475-487 pdf


16. Mokkonen M., Koskela E., Mappes T. & Mills S.C. 2008. The role of testosterone in physiology and evolution of a mammalian species. Acta Physiologica 193: S1604.

15. Poikonen, T., Koskela, E., Mappes T. & Mills S.C. 2008 Infanticide in the evolution of reproductive synchrony: effects on reproductive success. Evolution 62: 612-621 pdf

14. Mills S.C., Hazard, L., Lancaster, L., Mappes T., Miles, D., Oksanen, T.A. and Sinervo, B. 2008 Gonadotropin hormone modulation of testosterone, immune function, performance and behavioural tradeoffs among male morphs of the lizard, Uta stansburiana American Naturalist 171: 339-357 pdf


13. Mills S.C., Alatalo, R.V., Koskela, E., Mappes, J., Mappes T. & Oksanen T.A. 2007 Signal reliability compromised by genotype by environment interaction and potential mechanisms for its preservation. Evolution 61-7: 1748-1757 pdf

12. Mills S.C., Grapputo, A., Koskela, E. & Mappes T. 2007 Quantitative measure of sexual selection with respect to the operational sex ratio: a comparison of selection indices. Proceedings of the Royal Society of London B. 274 (1606): 143-150 pdf


11. Radwan J., Chadziska, M., Cicho, M., Mills, S.C., Matua, B., Sadowska, E.T., Baliga, K., Stanisz, A., Opuch, S. & Kotja, P. 2006 Metabolic costs of sexual advertisement in the bank vole Clethrionomys glareolus. Evolutionary Ecology Research 8: 859-869 pdf


10. Mills S.C., Taylor, M. & Reynolds, J.D. 2005 Benefits and costs to mussels from ejecting bitterling embryos: a test of the evolutionary equilibrium hypothesis. Animal Behaviour 70: 31-37 pdf


9. Mills S.C., & Reynolds, J.D. 2004. The importance of species interactions in conservation: the endangered European bitterling, Rhodeus sericeus and its freshwater mussel hosts. Animal Conservation 7(3): 257-263 pdf


8. Mills S.C., & Reynolds, J.D. 2003. Operational sex ratio and alternative reproductive behaviours in the European bitterling, Rhodeus sericeus. Behavioural Ecology and Sociobiology 54: 98-104 pdf

7. Mills S.C., & Reynolds, J.D. 2003. The bitterling-mussel interaction as a test case for co- evolution Journal of Fish Biology 63 SA: 84-104 pdf

6. Harper, D.M., Childress, R.B., Harper, M.M., Boar, R.R., Hickley, P., Mills S.C., Otieno, N., Drane, P., Vareschi, E, Nasirwa, O., Mwatha, W.E., Darlington, J.P., & Escuté-Gasulla, E.C. 2003. Aquatic biodiversity and saline lakes: Lake Bogoria National Reserve, Kenya. Hydrobiologia 500: 259-276 pdf

5. Mills S.C., & Côté, I. 2003. Sex-related differences in the growth and morphology of the blue mussel, Mytilus edulis. Journal of the Marine Biological Association 83: 1053-1057 pdf


4. Mills S.C., & Reynolds, J.D. 2002. Host species preferences by bitterling, Rhodeus sericeus, spawning in freshwater mussels and consequences for offspring survival. Animal Behaviour 63: 1029-1036 pdf

3. Mills S.C., & Reynolds, J.D. 2002. Mussel ventilation rates as a proximate cue for host selection by bitterling, Rhodeus sericeus. Oecologia 131: 473-478 pdf


2. Mills S.C., Peyrot-Clausade, M., & Fontaine, M.-F. 2000 Ingestion and transformation of algal turf by Echinometra mathaei on Tiahura fringing reef (French Polynesia). Journal of Experimental Marine Biology and Ecology 254: 71-84 pdf


1. Jennings, S., Reynolds, J.D., & Mills S.C. 1998 Life history correlates of responses to fisheries exploitation. Proceedings of the Royal Society of London B 265: 333-339 pdf



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